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In ruminants, cellulose is first hydrolysed by ruminal microorganisms into cellobiose, then is fermented to pyruvate and finally volatile fatty acids. The changes of forage to concentrate ratios in the diet significantly affect the number and type of rumen microorganisms and then affect the end products of fermentation. Moreover, the extent of cellulose digestion is a compromise between the rate of hydrolysis and the retention time in the rumen related to the particle size of the forage. The intrinsic digestibility of cellulose depends on the origin and treatment of the forage. As far as cellulose is associated to lignin, hemicelluloses and cutin in natural forages, a wide range of digestibility is observed (30 to 90%). Crystallinity of cellulose decreases the rate but not the extent of digestibility that may reach 80% (Van Soest, 1994).